Ben Stiller

Alex (voice)

Chris Rock

Marty (voice)

David Schwimmer

Melman (voice)

Jada Pinkett Smith

Gloria (voice)

Frances McDormand

Captain Chantel DuBois (voice)

Sacha Baron Cohen

Julien (voice)

Andy Richter

Mort (voice)

Cedric the Entertainer

Maurice (voice)

Jessica Chastain

Gia (voice)

Bryan Cranston

Vitaly (voice)

Martin Short

Stefano (voice)

Paz Vega

Carmen (voice)

Frank Welker

Sonya (voice)

Tom McGrath

Skipper / First Policeman (voice)

Chris Miller

Kowalski (voice)

Christopher Knights

Private (voice)

Conrad Vernon

Mason / Second Policeman (voice)

Vinnie Jones

Freddie the Dog (voice)

Steve Jones

Jonesy the Dog (voice)

Nick Fletcher

Frankie the Dog (voice)

Danny Jacobs

Croupier / Circus Master (voice)

Dan O'Connor

Casino Security / Mayor of New York City (voice)

Eric Darnell

Comandante / Zoo Official / Zoo Announcer (voice)

Stephen Kearin

Fourth Policeman (voice)

Emily Nordwind

Zoo Official (voice)

Asucena Jimenez-Cruz

Kid in Crowd (voice)